Zhang et. al. published the research on structure and properties of honeybee combs on PNAS
PNAS published the paper of Hierarchical, multilayered cell walls reinforced by recycled silk cocoons enhance the structural integrity of honeybee combs (Proc. Natl. Acad. Sci. USA, 107 (21): 9502—9506) by Kai Zhang, Huiling Duan and Jianxiang Wang cooperating with Bhushan Karihaloo from the College of Engineering of Cardiff University.